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Oli, M.K. |
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Title |
Seasonal patterns in habitat use of blue sheep Pseudois nayaur (Artiodactyla, Bovidae) in Nepal |
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1996 |
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Mammalia |
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60 |
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2 |
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187-193 |
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blue-sheep; snow-leopard; Panthera-uncia; Nepal; conservation; prey; predator; snow leopard; blue; sheep; browse; panthera; uncia; 670 |
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Blue sheep (Pseudois nayaur) are the main prey of the endangered snow leopard (Panthera uncia) as well as an important game species in Nepal. A knowledge of how blue sheep utilize their habitat is essential for the scientific management of the sheep and for the conservation of the snow leopard, but we only have a limited understanding of this aspect of blue sheep ecology. I studied the habitat use pattern of blue sheep by direct observation in the Anna-purna Conservation Area, Nepal where they occur sympatrically with the snow leopard. The sheep used grassland habitats more frequently during pre-parturition (spring) and post-parturition (autumn) than other habitat types, but scrub and grassland habitats were used equally frequently during the rut (winter). The sheep used smooth undulating slopes of medium steepness (<40 degrees) on southerly aspects within the elevation range of 4,200-4,600 m most frequently in all seasons, and there was no evidence of seasonal migration along the elevation gradient. When not in broken landforms (e.g., cliff, landslides), the sheep maintained proximity (less than or equal to 150 m) to such features suggesting their importance as escape cover (i.e., from predators). The use of habitat components by blue sheep appeared to be related to the distribution of foraging areas and escape cover. |
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UNIV EDINBURGH,INST CELL ANIM & POPULAT BIOL,EDINBURGH EH9 3JT,MIDLOTHIAN,SCOTLANDANNAPURNA CONSERVAT AREA PROJECT,KATMANDU,NEPAL /Publisher:MUSEUM NAT HIST NATURELLE, PARIS Document Type: English |
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SLN @ rana @ 289 |
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751 |
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Oli, M.K.; Taylor, K.R.; Rogers, M.E. |
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Title |
Snow leopard Panthera uncia predation of livestock: An assessment of local perceptions in the Annapurna Conservation Area, Nepal |
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Miscellaneous |
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Year |
1994 |
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Biological Conservation |
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68 |
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63-68 |
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snow leopard,blue sheep,livestock predation,public attitudes,Annapurna Conservation Area,Nepal. |
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Public attitudes towards snow leopard Panthera uncia predation of domestic livestock were investigated by a questionnaire survey of four villages in snow leopard habitat within the Annapurna Conservation Area, Nepal Most local inhabitants were subsistence farmers, many dependent upon yaks, oxen, horses and goats, with an average livestock holding of 26.6 animals per household. Reported losses to snow leopards averaged 0.6 and O. 7 animals per household in two years of study, constituting 2.6% of total stockholding but representing in monetary terms almost a quarter of the average annual Nepali national per capita income. Local people hem strongly negative attitudes towards snow leopards and most suggested that total extermination of leopards was the only acceptable solution to the predation problem. Snow leopards were reported to be killed by herdsmen in defence of their livestock. The long-term success of snow leopard conservation programmes may depend upon the satisfactory resolution of the predation conflict. Some possible ways of reducing predation losses are also discussed. |
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SLN @ rana @ 893 |
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753 |
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Prasad, S.N.; Chundawat, R.S.; Hunter, D.O.; Panwar, H.S.; Rawat, G.S. |
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Remote sensing snow leopard habitat in the trans-Himalaya of India using spatial models and satellite imagery preliminary results |
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1991 |
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519-523 |
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snow-leopard; Gis; cartographic-modelling; India; Ladakh; Zanskar; predation; habitat; prey; predator; blue-sheep; snow leopard; blue; sheep; browse; cartographic modelling; cartographic; modelling; 810 |
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The snow leopard (Panthera uncia) is a flagship species for conservation in the high mountain regions of central Asia. Data on snow leopard predation, habitat conditions and range of main prey species were gathered along with thematic maps of the study area for elevation, snow cover, sighting data, kill data, blue sheep use areas, and vegetation data. These data were entered into a GIS and used to help delineate surface features from a satellite image. Preliminary results show that general physiographic features of snow leopard habitat can be detected using satellite imagery and that GIS cartographic modeling techniques can improve this delineation. -from Authors |
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Buhyoff, G.J. |
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Title, Monographic: Resource Technology 90. Proc. second international symposium on advanced technology in natural resources management
Place of Meeting: Washington, DC |
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SLN @ rana @ 176 |
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792 |
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Mishra, C.; Van Wieren S.; Ketner, P.; Heitkonig, I.; Prins H. |
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Competition between domestic livestock and wild bharal Pseudois nayaur in the Indian Trans-Himalaya |
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2004 |
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Journal of Animal Ecology |
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73 |
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344-354 |
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blue sheep,diet,grazing,rangeland,resource,ungulate,wildlife; 5320 |
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1. The issue of competition between livestock and wild herbivores has remained contentious. We studied the diets and population structures of the mountain ungulate bharal Pseudois nayaur and seven species of livestock to evaluate whether or not they compete for forage. The study was conducted in the high altitude Spiti Valley, Indian Trans-Himalaya.
2. We compared resource (forage) availability and bharal population structures between rangelands differing in livestock density. Forage availability was estimated by clipping the standing graminoid biomass in sample plots. Livestock and bharal population structures were quantified through annual censuses. Seasonal diets of livestock were studied by direct observations, while those of bharal were quantified through feeding
signs on vegetation.
3. We found that livestock grazing causes a significant reduction in the standing crop of forage. Graminoid availability per unit livestock biomass was three times greater in a moderately grazed rangeland compared with an intensively grazed one.
4. There was considerable diet overlap among the herbivore species. In summer, bharal, yak Bos grunniens, horse Equus caballus, cow Bos indicus, and dzomo (yak-cow hybrids) fed predominantly on graminoids, while donkey E. asinus, sheep Ovis aries, and goat Capra hircus, consumed both graminoids and herbs. The summer diet of bharal was a subset of the diets of three livestock species. In winter, depleted graminoid availability caused bharal, yak and horse to consume relatively more herbs, while the remaining livestock species fed predominantly on graminoids. Diet overlap was less in winter but, in both seasons, all important forage species in the bharal diet were consumed
in substantial amounts by one or more species of livestock.
5. Comparison of the population structures of bharal between two rangelands differing in livestock density by
c. 30% yielded evidence of resource competition. In the intensively grazed rangeland, bharal density was 63% lower, and bharal population showed poorer performance (lower young : adult female ratios).
6.Synthesis and applications High diet overlap between livestock and bharal, together with density-dependent forage limitation, results in resource competition and a decline in bharal density. Under the present conditions of high livestock density and supplemental feeding, restricting livestock numbers and creating livestockfree areas are necessary measures for conserving Trans-Himalayan wild herbivores. Mediating competitive effects on bharal through supplemental feeding is not a feasible option. |
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SLN @ rana @ 517 |
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695 |
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Author |
Schaller, G.B. |
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Title |
On the behaviour of Blue Sheep (Pseudois nayaur) |
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1972 |
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Journal of Bombay Natural Historical Society |
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69 |
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523-537 |
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predator; prey; hunting; scat; blue-sheep; Nepal; blue; sheep; browse; 2230 |
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Two or three snow leopards hunted in the study area in eastern Nepal. Describes content of some snow leopard scat |
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SLN @ rana @ 24 |
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862 |
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Oli, M. |
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Title |
Snow leopards and blue sheep in Nepal: Densities and predator: Prey ratio |
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Miscellaneous |
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1994 |
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Journal of Mammalogy |
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75 |
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998-1004 |
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snow leopard,Panthera uncia,blue sheep,Pseudois nayaur,density,predator:prey ratio,harvest rate,livestock predation,Nepal |
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I studied snow leopards (Panthera uncia) and blue sheep (Pseudois nayaur) in Manang District, Annapurna Conservation Area, Nepal, to estimate numbers and analyze predatorprey interactions. Five to seven adult leopards used the 105-km2 study area, a density of 4.8 to 6.7 leopards/100 km2. Density of blue sheep was 6.6-10.2 sheep/km2, and biomass density was 304 kg/km2. Estimated relative biomass consumed by snow leopards suggested that blue sheep were the most important prey; marmots (Marmota himalayana) also contributed significantly to the diet of snow leopards. Snow leopards in Manang were estimated to harvest 9-20% of total biomass and 11-24% of total number of blue sheep annually. Snow leopard :blue sheep ratio was 1 :1 14-1 :159 on a weight basis, which was considered sustainable given the importance of small mammals in the leopard's diet and the absence of other competing predators. |
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SLN @ rana @ 894 |
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741 |
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Schaller, G.B.; Jurang, R.; Mingjiang, Q. |
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Status of snow leopard (Panthera-uncia) in Qinghai-Province and Gansu Province-China |
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1988 |
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Biological Conservation |
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45 |
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3 |
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179-194 |
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status; population; China; Qinhai-province; Gansu-province; conservation; snow-leopard; Panthera-uncia; prey; ungulates; blue-sheep; marmot; snow leopard; blue; sheep; browse; qinhai province; qinhai; province; gansu province; gansu; panthera uncia; panthera; uncia; 860 |
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The status and distribution of the snow leopard Panthera uncia was investigated in two provinces of China. The cats occur over about 65,000km2 or 9% of the Qinghai Province, and in a few places along the western edge of Gansu Province. In many areas the animals have in recent decades been decimated or locally eradicated, as have their prey. Counts of wild ungulates in 9 mountain block, totalling 1375km2, known for abundant wildlife, had an average of 1.4-5.4 animals km2, principally blue sheep Psuedois nayaur, which together with marmot Marmota himalayana, represent the snow leopards main prey. Possibly 650 snow leopards survive in Qinghai but shooting and trapping of this legally protected animal and the hunting of blue sheep for local consumtion and export threaten their existence. |
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Publisher:ELSEVIER SCI LTD, OXFORD |
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SLN @ rana @ 113 |
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867 |
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Shrestha, B. |
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Prey Abundance and Prey Selection by Snow Leopard (uncia uncia) in the Sagarmatha (Mt. Everest) National Park, Nepal |
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2008 |
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1-35 |
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project; snow; snow leopard; snow-leopard; leopard; network; conservation; program; prey; abundance; selection; uncia; Uncia uncia; Uncia-uncia; Sagarmatha; national; national park; National-park; park; Nepal; resource; predators; predator; ecological; impact; region; community; structure; number; research; population; status; density; densities; wild; prey species; prey-species; species; Himalayan; tahr; musk; musk-deer; deer; game; birds; diet; livestock; livestock depredation; livestock-depredation; depredation; awareness; co-existence; ungulates; ungulate; Human; using; areas; area; monitoring; transect; Hair; identification; scat; attack; patterns; sighting; 1760; populations; birth; Male; Female; young; domestic; domestic livestock; 120; scats; yak; Dog; pika; wildlife; Seasons; winter; horse; study; cover; land; predation; Pressure; development; strategy; threatened; threatened species; threatened-species; conflicts; conflict; people; control; husbandry; compensation; reintroduction; blue; blue sheep; blue-sheep; sheep; free ranging |
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Predators have significant ecological impacts on the region's prey-predator dynamic and community structure through their numbers and prey selection. During April-December 2007, I conducted a research in Sagarmatha (Mt. Everest) National Park (SNP) to: i) explore population status and density of wild prey species; Himalayan tahr, musk deer and game birds, ii) investigate diet of the snow leopard and to estimate prey selection by snow leopard, iii) identify the pattern of livestock depredation by snow leopard, its mitigation, and raise awareness through outreach program, and identify the challenge and opportunities on conservation snow leopard and its co-existence with wild ungulates and the human using the areas of the SNP. Methodology of my research included vantage points and regular monitoring from trails for Himalayan tahr, fixed line transect with belt drive method for musk deer and game birds, and microscopic hair identification in snow leopard's scat to investigate diet of snow leopard and to estimate prey selection. Based on available evidence and witness accounts of snow leopard attack on livestock, the patterns of livestock depredation were assessed. I obtained 201 sighting of Himalayan tahr (1760 individuals) and estimated 293 populations in post-parturient period (April-June), 394 in birth period (July -October) and 195 November- December) in rutting period. In average, ratio of male to females was ranged from 0.34 to 0.79 and ratio of kid to female was 0.21-0.35, and yearling to kid was 0.21- 0.47. The encounter rate for musk deer was 1.06 and density was 17.28/km2. For Himalayan monal, the encounter rate was 2.14 and density was 35.66/km2. I obtained 12 sighting of snow cock comprising 69 individual in Gokyo. The ratio of male to female was 1.18 and young to female was 2.18. Twelve species (8 species of wild and 4 species of domestic livestock) were identified in the 120 snow leopard scats examined. In average, snow leopard predated most frequently on Himalayan tahr and it was detected in 26.5% relative frequency of occurrence while occurred in 36.66% of all scats, then it was followed by musk deer (19.87%), yak (12.65%), cow (12.04%), dog (10.24%), unidentified mammal (3.61%), woolly hare (3.01%), rat sp. (2.4%), unidentified bird sp. (1.8%), pika (1.2%), and shrew (0.6%) (Table 5.8 ). Wild species were present in 58.99% of scats whereas domestic livestock with dog were present in 40.95% of scats. Snow leopard predated most frequently on wildlife species in three seasons; spring (61.62%), autumn (61.11%) and winter (65.51%), and most frequently on domestic species including dog in summer season (54.54%). In term of relative biomass consumed, in average, Himalayan tahr was the most important prey species contributed 26.27% of the biomass consumed. This was followed by yak (22.13%), cow (21.06%), musk deer (11.32%), horse (10.53%), wooly hare (1.09%), rat (0.29%), pika (0.14%) and shrew (0.07%). In average, domestic livestock including dog were contributed more biomass in the diet of snow leopard comprising 60.8% of the biomass consumed whilst the wild life species comprising 39.19%. The annual prey consumption by a snow leopard (based on 2 kg/day) was estimated to be three Himalayan tahr, seven musk deer, five wooly hare, four rat sp., two pika, one shrew and four livestock. In the present study, the highest frequency of attack was found during April to June and lowest to July to November. The day of rainy and cloudy was the more vulnerable to livestock depredation. Snow leopard attacks occurred were the highest at near escape cover such as shrub land and cliff. Both predation pressure on tahr and that on livestock suggest that the development of effective conservation strategies for two threatened species (predator and prey) depends on resolving conflicts between people and predators. Recently, direct control of free – ranging livestock, good husbandry and compensation to shepherds may reduce snow leopard – human conflict. In long term solution, the reintroduction of blue sheep at the higher altitudes could also “buffer” predation on livestock. |
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Project funded by Snow Leopard Network's Snow Leopard Conservation Grant Program. Forum of Natural Resource Managers, Nepal. |
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SLN @ rana @ 1076 |
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887 |
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Shrestha, R.; Wegge, P.; Koirala, R.A. |
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Summer diets of wild and domestic ungulates in Nepal Himalaya |
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2005 |
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Journal of Zoology |
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266 |
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111-119 |
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argali; blue sheep; Capra hircus; diet; faecal analysis; fecal analysis; mountain ungulates; Ovis ammon hodgsoni; pseudois nayaur |
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The selection of summer forage by three sympatric ungulates in the Damodar Kunda region of upper Mustang in
north Nepal was studied to assess the extent of food overlap between them. To compare their diets, a microhistological technique of faecal analysis was used, adjusted for inherent biases by comparing it with bite-count data obtained in domestic goats. Tibetan argali Ovis ammon hodgsoni, naur (blue sheep or bharal) Pseudois nayaur and domestic goat Capra hircus consumed mostly forbs, graminoids and browse, respectively. The proportions of food items in their diets were significantly different both at the plant species (P<0.02) and at the forage category level (P<0.001). Except for sharing three common plants (Agrostis sp., Stipa sp. and Potentilla fruticosa), dietary overlap at the species level was quite low. At the forage category level, naur and domestic goat overlapped more than the other ungulate pairs. Although all three species were opportunistic, mixed feeders, argali was a more selective forb specialist grazer than the other two ungulates. Owing to some spatial separation and little dietary overlap, interspecific competition for summer forage was low. If animal densities increase, however, goats are expected to compete more with naur than with argali because of their more similar diets. Owing to differences in forage selection by argali and naur throughout their large geographical ranges, reflecting adaptations to local ecological conditions, inferences regarding forage competition between domestic livestock and these two wild caprins need to be made from local, site-specific studies, rather than from general diet comparisons. |
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The Zoological Society of London |
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London |
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SLN @ rana @ 934 |
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888 |
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Shrestha, R.; Wegge, P. |
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Wild sheep and livestock in Nepal Trans-Himalaya: coexistence or competition? |
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2008 |
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Environmental Conservation |
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32 |
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2 |
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125-136 |
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bharal; blue sheep; diet; food habits; mountain ungulates; Nepal; pastoralism; pseudois nayaur; rangeland management; resource partitioning |
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Excessive grazing by livestock is claimed to displace wild ungulates in the Trans-Himalaya. This study compares the seasonal diets and habitat use of sympatric wild naur Pseudois nayaur and domestic goat Capra hircus, sheep Ovis aries and free-ranging yak Bos grunniens in north Nepal and analyses their overlap both within and across seasons. Alpinemeadow and the legumes Oxytropis and Chesneya were critical resources for all animal groups. High overlap occurred cross-seasonally when smallstock (sheep and goats) in summer used the spring and autumn ranges of naur. Relatively high total ungulate biomass (3028 kg km-2) and low recruitment of naur (56 young per 100 adult females in autumn) suggested interspecific competition. The spatio-temporal heterogeneity in composition and phenology of food plants across the steep gradient of altitude, together with rotational grazing, appears to indirectly facilitate coexistence of naur and smallstock. However, owing to high crossseasonal (inter-seasonal) overlaps, competition is likely to occur between these two groups at high stocking densities. Within seasons, naur overlapped more with free-ranging yak than with smallstock. As their habitat use and diets were most similar in winter, when both fed extensively on the same species of shrubs, naur was most likely to compete with yak during that season.
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SLN @ rana @ 936 |
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890 |
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