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Saltz, D., Rowen, M., & Rubenstein, D. (2000). The effect of space-use patterns of reintroduced Asiatic wild ass on effective population size. Conservation Biology, 14(6), 1852–1861.
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Mishra, C., Van Wieren S., Ketner, P., Heitkonig, I., & Prins H. (2004). Competition between domestic livestock and wild bharal Pseudois nayaur in the Indian Trans-Himalaya. Journal of Animal Ecology, 73, 344–354.
Abstract: 1. The issue of competition between livestock and wild herbivores has remained contentious. We studied the diets and population structures of the mountain ungulate bharal Pseudois nayaur and seven species of livestock to evaluate whether or not they compete for forage. The study was conducted in the high altitude Spiti Valley, Indian Trans-Himalaya.
2. We compared resource (forage) availability and bharal population structures between rangelands differing in livestock density. Forage availability was estimated by clipping the standing graminoid biomass in sample plots. Livestock and bharal population structures were quantified through annual censuses. Seasonal diets of livestock were studied by direct observations, while those of bharal were quantified through feeding
signs on vegetation.
3. We found that livestock grazing causes a significant reduction in the standing crop of forage. Graminoid availability per unit livestock biomass was three times greater in a moderately grazed rangeland compared with an intensively grazed one.
4. There was considerable diet overlap among the herbivore species. In summer, bharal, yak Bos grunniens, horse Equus caballus, cow Bos indicus, and dzomo (yak-cow hybrids) fed predominantly on graminoids, while donkey E. asinus, sheep Ovis aries, and goat Capra hircus, consumed both graminoids and herbs. The summer diet of bharal was a subset of the diets of three livestock species. In winter, depleted graminoid availability caused bharal, yak and horse to consume relatively more herbs, while the remaining livestock species fed predominantly on graminoids. Diet overlap was less in winter but, in both seasons, all important forage species in the bharal diet were consumed
in substantial amounts by one or more species of livestock.
5. Comparison of the population structures of bharal between two rangelands differing in livestock density by
c. 30% yielded evidence of resource competition. In the intensively grazed rangeland, bharal density was 63% lower, and bharal population showed poorer performance (lower young : adult female ratios).
6.Synthesis and applications High diet overlap between livestock and bharal, together with density-dependent forage limitation, results in resource competition and a decline in bharal density. Under the present conditions of high livestock density and supplemental feeding, restricting livestock numbers and creating livestockfree areas are necessary measures for conserving Trans-Himalayan wild herbivores. Mediating competitive effects on bharal through supplemental feeding is not a feasible option.
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Filonov K.F. (1996). Large terrestrial mammals in the reserves of Russia: their status and prospects of conservation.
Abstract: The authors make an analysis of fauna of large mammals in 68 nature reserves. There are 10 carnivores and 17 ungulates. Wolf, brown bear, wolverine and lynx appeared to be more widely spread. Dhole, snow leopard, tiger, Himalayan bear have limited distribution and low density. Hey have recorded in a few nature reserves. Among the ungulates wild boar, musk deer, red deer, roe deer, moose, reindeer and aurochs are more widely spread.
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Miller, D. J., & Jackson, R. (1994). Livestock and Snow Leopards:making room for competing users on the Tibetian Plateau. In J.L.Fox, & D.Jizeng (Eds.), (pp. 315–328). Usa: Islt.
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Yongsheng, L. (1994). International hunting and the involvement of local people in Dulan, Qinghai, China. In J.L.Fox, & D.Jizeng (Eds.), (pp. 305–314). Usa: Islt.
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Zhiryakov V.A. (1990). Wolves' role in biocenosis of the Almaty nature reserve (North Tien Shan) (Vol. Vol. II.).
Abstract: The quantity of ungulates is high in the nature reserve: moral (100-120), roe deer (500-650), Siberian ibex (660-700), and wild boar (50-80). Moreover some 5,000 heads of livestock (mostly sheep) are grazed in a buffer zone in summer. Among big predators (snow leopard, bear, lynx) wolf kills about 40 percent of ungulates.
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Ismagilov M.I. (1983). Protection of rare mammals in Kazakhstan.
Abstract: The following rare mammals can be found in nature reserves of Kazakhstan: argali, goitered gazelle, kulan, snow leopard, stone marten, Tien Shan brown bear, manul, Turkistan lynx, Menzbier's marmot, and porcupine. The rest of rare mammal species (three insectivorous species, seven rodent, eight predator, and two ungulate species) are outside of protected areas and require special protection measures.
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Braden, K. E. (1988). Economic Development in Six Regions of Snow Leopard Habitat in the U.S.S.R. In H.Freeman (Ed.), (pp. 227–246). India: International Snow Leopard Trust and the Wildlife Institute of India.
Abstract: The Disappearance of traditional ungulate prey of the snow leopard may be contributing to its endangered status in the wild. Soviet biologists have noted that wild sheep are a primary prey of the snow leopard in the southern Russian union republic and the Central Asian union republic of the U.S.S.R. While poaching appears to have had some impact on the status of these sheep, econmic pressures may be contributing to their decrease. Evidence presented for KAzakhstan and three regions of the Russian republic demonstrates that commercial sheep and goat production appears to be growing at a very high pace in these areas, thus consumming habitat otherwise available for wild herds.
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Zhiryakov V.A. (1989). The influence of the predators on population trend of the ungulates in the Almaty nature reserve.
Abstract: The data on predators and ungulates population dynamics in Almaty Nature reserve (Kazakhstan) in 1983-1987s are given. The number of snow leopard is stable (3-5 individuals), the density is 0.06 indi/1000 ha. An insignificant increase of Siberian ibex' number (660 to 700) with density of 36 indi/1000 ha is recorded.
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Schaller, G. B., Jurang, R., & Mingjiang, Q. (1988). Status of snow leopard (Panthera-uncia) in Qinghai-Province and Gansu Province-China. Biological Conservation, 45(3), 179–194.
Abstract: The status and distribution of the snow leopard Panthera uncia was investigated in two provinces of China. The cats occur over about 65,000km2 or 9% of the Qinghai Province, and in a few places along the western edge of Gansu Province. In many areas the animals have in recent decades been decimated or locally eradicated, as have their prey. Counts of wild ungulates in 9 mountain block, totalling 1375km2, known for abundant wildlife, had an average of 1.4-5.4 animals km2, principally blue sheep Psuedois nayaur, which together with marmot Marmota himalayana, represent the snow leopards main prey. Possibly 650 snow leopards survive in Qinghai but shooting and trapping of this legally protected animal and the hunting of blue sheep for local consumtion and export threaten their existence.
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Fox, J. L., Nurbu, C., & Chundawat, R. S. (1991). The Mountain Ungulates of Ladakh India. Biological Conservation, 58, 167–190.
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Zhiryakov V.A. (1976). Ibex. Rare ungulate species of the Almaty nature reserve and their protection.
Abstract: Collected are data on rare ungulates in the Almaty nature reserve in 1968-1973. Since recently the population of goitered gazelle has dropped sharply and is now 20-30 animals per seven ha. The nature reserve shall be expanded in order to protect the animals. Argali inhabits a desert area in the mountains of Greater and Lesser Kalkana. Argali sometimes migrates outside the nature reserve. Ibex inhabits a mountainous part of the nature reserve, its population being 10-13 animals per 1,000 ha. Predators have negligible impact on the ibex population (12.5 percent of deaths), which is preyed on solely by snow leopard and wolf.
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Shrestha, R., & Wegge, P. (2008). Wild sheep and livestock in Nepal Trans-Himalaya: coexistence or competition? Environmental Conservation, 32(2), 125–136.
Abstract: Excessive grazing by livestock is claimed to displace wild ungulates in the Trans-Himalaya. This study compares the seasonal diets and habitat use of sympatric wild naur Pseudois nayaur and domestic goat Capra hircus, sheep Ovis aries and free-ranging yak Bos grunniens in north Nepal and analyses their overlap both within and across seasons. Alpinemeadow and the legumes Oxytropis and Chesneya were critical resources for all animal groups. High overlap occurred cross-seasonally when smallstock (sheep and goats) in summer used the spring and autumn ranges of naur. Relatively high total ungulate biomass (3028 kg km-2) and low recruitment of naur (56 young per 100 adult females in autumn) suggested interspecific competition. The spatio-temporal heterogeneity in composition and phenology of food plants across the steep gradient of altitude, together with rotational grazing, appears to indirectly facilitate coexistence of naur and smallstock. However, owing to high crossseasonal (inter-seasonal) overlaps, competition is likely to occur between these two groups at high stocking densities. Within seasons, naur overlapped more with free-ranging yak than with smallstock. As their habitat use and diets were most similar in winter, when both fed extensively on the same species of shrubs, naur was most likely to compete with yak during that season.
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Sukhbat, K., & Munkhtsog, B. (1997). Density and Distribution of Ibex and Argali Sheep in Mongolia. In R.Jackson, & A.Ahmad (Eds.), (pp. 121–123). Lahore, Pakistan: Islt.
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Shrestha, R., Wegge, P., & Koirala, R. A. (2005). Summer diets of wild and domestic ungulates in Nepal Himalaya. Journal of Zoology, 266, 111–119.
Abstract: The selection of summer forage by three sympatric ungulates in the Damodar Kunda region of upper Mustang in
north Nepal was studied to assess the extent of food overlap between them. To compare their diets, a microhistological technique of faecal analysis was used, adjusted for inherent biases by comparing it with bite-count data obtained in domestic goats. Tibetan argali Ovis ammon hodgsoni, naur (blue sheep or bharal) Pseudois nayaur and domestic goat Capra hircus consumed mostly forbs, graminoids and browse, respectively. The proportions of food items in their diets were significantly different both at the plant species (P<0.02) and at the forage category level (P<0.001). Except for sharing three common plants (Agrostis sp., Stipa sp. and Potentilla fruticosa), dietary overlap at the species level was quite low. At the forage category level, naur and domestic goat overlapped more than the other ungulate pairs. Although all three species were opportunistic, mixed feeders, argali was a more selective forb specialist grazer than the other two ungulates. Owing to some spatial separation and little dietary overlap, interspecific competition for summer forage was low. If animal densities increase, however, goats are expected to compete more with naur than with argali because of their more similar diets. Owing to differences in forage selection by argali and naur throughout their large geographical ranges, reflecting adaptations to local ecological conditions, inferences regarding forage competition between domestic livestock and these two wild caprins need to be made from local, site-specific studies, rather than from general diet comparisons.
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Dementiev G.P. (1967). Quadrupeds inhabitants of the mountains.
Abstract: All species inhabiting the highlands of Asia are normally referred to as herbivorous or predators. A majority of alpine land species (rodents and ungulates) feeds upon leaves, stalks, and roots of plants. Among widely distributed highland species the most interesting are marmots, red pica, grey vole, argali, and ibex. Argali and ibex are preyed on by snow leopards. There are reasons to believe that these mountain animal species are more ancient than their cognates in a plain. All the way from Central Asia to Europe, species belonging to the eastern and western fauna complexes are observed to interpenetrate.
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Harris, R. B. (1994). Dealing with uncertainty in counts of mountain ungulates. In J.L.Fox, & D. Jizeng (Eds.), (pp. 105–111). Usa: Islt.
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Kovshar A.F. (1982). Preservation of gene pool of rare and endangered animal species.
Abstract: The rare species are protected in six nature reserves in Kazakhstan, including 9 mammals, 29 birds, and one reptile species. More than 20 rare and endangered species inhabiting Kazakhstan cannot be met within the nature reserves. The point is to establish a network of state nature reserves, particularly in steppe and desert area of the country.
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Kohli, K., Sankaran, M., Suryawanshi, K. R., Mishra, C. (2014). A penny saved is a penny earned: lean season foraging strategy of an alpine ungulate. Animal Behaviour, (92), 93–100.
Abstract: Lean season foraging strategies are critical for the survival of species inhabiting highly seasonal environments
such as alpine regions. However, inferring foraging strategies is often difficult because of
challenges associated with empirically estimating energetic costs and gains of foraging in the field. We
generated qualitative predictions for the relationship between daily winter foraging time, body size and
forage availability for three contrasting foraging strategies including time minimization, energy intake
maximization and net energy maximization. Our model predicts that for animals employing a time
minimization strategy, daily winter foraging time should not change with body size and should increase
with a reduction in forage availability. For energy intake maximization, foraging time should not vary
with either body size or forage availability. In contrast, for a net energy maximization strategy, foraging
time should decrease with increase in body size and with a reduction in forage availability. We contrasted
proportion of daily time spent foraging by bharal, Pseudois nayaur, a dimorphic grazer, across
different body size classes in two high-altitude sites differing in forage availability. Our results indicate
that bharal behave as net energy maximizers during winter. As predicted by the net energy maximization
strategy, daily winter foraging time of bharal declined with increasing body size, and was lower in the
site with low forage availability. Furthermore, as predicted by our model, foraging time declined as the
winter season progressed. We did not find support for the time minimizing or energy intake maximizing
strategies. Our qualitative model uses relative rather than absolute costs and gains of foraging which are
often difficult to estimate in the field. It thus offers a simple way to make informed inferences regarding
animal foraging strategies by contrasting estimates of daily foraging time across gradients of body size
and forage availability.
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Heiz A.V. (1983). Snow leopard in Kyrgyzstan and its protection (Vol. 3).
Abstract: In the year 1970, the quantity of snow leopards in Kyrgyzstan was defined as 1,300 animals, while in the years to follow 1,600 animals were recorded. A snow leopard population has significantly decreased since recently because of intense extermination of snow leopard's prey ungulates, particularly ibex. In some areas of the Kyrgyz ridge livestock is growing in number thus affecting snow leopard population. It is extremely rare that snow leopard would attack livestock. Snow leopards can be caught under special license. Educational and awareness work among shepherds and hunters residing in the mountainous area of the country needs to be improved.
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Ale, S. B., Yonzon, P., & Thapa, K. (2007). Recovery of snow leopard Uncia uncia in Sagarmatha (Mount Everest) National Park, Nepal (Vol. 41).
Abstract: From September to November 2004 we conducted surveys of snow leopard Uncia uncia signs in three major valleys in Sagarmatha (Mount Everest) National Park in Nepal using the Snow Leopard Information Management System, a standardized survey technique for snow leopard research. We walked 24 transects covering c. 14 km and located 33 sites with 56 snow leopard signs, and 17 signs incidentally in other areas. Snow leopards appear to have re-inhabited the Park, following their disappearance c. 40 years ago, apparently following the recovery of Himalayan tahr Hemitragus jemlahicus and musk deer Moschus chrysogaster populations. Taken together the locations of all 73 recent snow leopard signs indicate that the species is using predominantly grazing land and shrubland/ open forest at elevations of 3,000-5,000 m, habitat types that are also used by domestic and wild ungulates. Sagarmatha is the homeland of c. 3,500 Buddhist Sherpas with .3,000 livestock. Along with tourism and associated developments in Sagarmatha, traditional land use practices could be used to ensure coexistence of livestock and wildlife, including the recovering snow leopards, and ensure the wellbeing of the Sherpas.
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Formozov A.N. (1990). Seasonal migrations of mammals due to snow cover. Distribution of the Felidae family species.
Abstract: It describes vertical migrations of ungulates (ibex, wild sheep) in the Semerechie, Altai, Sayans, Tuva, seasonal migrations of steppe ungulates (kulan and saiga), and migrations of predators (lynx, leopard, irbis, tiger, dhole, wolf, glutton) following ungulates during winters with thick snow cover. Shorter local migrations related to uneven snow cover are typical for corsac, fox, and wolf. An analysis of the Felidae family species distribution showed that northern border of the cat family species habitat is connected with borders of 20 30 cm thick snow cover rather than with landscape contours or typical habitats. With the exception of lynx, this can be referred to the large cat family species such as irbis, leopard, and tiger.
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Harris, R. B. (1994). A note on snow leopards and local people in Nangqian County, Southern Qinghai Province. In J.L.Fox, & D. Jizeng (Eds.), (pp. 79–84). Usa: Islt.
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Vorobjov A.G. (2002). Ungulates (Artiodactyla) of the Western Tien Shan (Distribution and number within Kyrgyzstan).
Abstract: Materials on numbers and densities of 8 ungulates (Sus scrofa nigripes, Capreolus pygargus tianschanicus, Cervus elaphus, Cervus nippon, Cervus dama, Capra sibirica formosovi, Ovis ammon karelini, Bison bonasus) within the Chatkal …nd Talas Ranges are given. A critical analysis is also presented on prospects for development of the commercial hunting tourism as opportunities for additional assignments in measures on conservation of the Western Tien Shan biodiversity
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Zhiryakov V.A. (1986). Snow leopard in the Almaty nature reserve. Short messages about snow leopards.
Abstract: Snow leopard is a common species for the Almaty nature reserve due to numerous wild ungulates, particularly ibexes (about 600 ibexes at a density of 32 animals per 1,000 ha) inhabiting the area. According to the data of 1982 there were 0.5 footprints of snow leopard per 10 km of transect. The remains of ibex, roe deer, squirrel, gray vole mouse and birds were found in faeces of snow leopards. Snow leopard attacks their prey unexpectedly, being in wait for it in such places where prey is difficult to escape from. When hunt is successful the prey is killed almost instantly. Snow leopard feeds upon the same prey for several days.
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