Singh, N. J. (2008). Animal – Habitat relationships in high altitude rangelands. Norway: University of Tromsø.
Abstract: This study conducted in the high altitude rangelands of Indian Transhimalaya, deals with basic questions regarding the ecology of an endangered species, the wildsheep Tibetan argali (Ovis ammon hodgsoni) and applied issues related to its conservation and potential conflict with the local nomadic pastoralists. The basic questions on ecology are aimed at delineating the habitat and resource selection processes, identifying factors causing sexual segregation and efficient surveying and sampling. The applied aspect focuses on the changing face of pastoralism and the potential impacts of modernising livestock husbandry on argali.
Overall, the study provides a general framework towards the understanding of argali-habitat relationships at different spatio-temporal scales. The spatial determinant associated with altitude in the area, predicts argali habitat and resource selection in this relatively homogenous landscape. These determine the range of other topographic variables and forage characteristics selected by argali. The selection of feeding patches in the selected range of altitude and topography is mainly characterised by their greenness and the quality of plant groups. Adjusting to changing forage quality, argali display an opportunistic feeding strategy, selecting grasses in early spring and switching to forbs later in summer. Nevertheless, the habitat selection process did not appear to differ among the sexes to drive sexual segregation. There was, however, strong segregation among the sexes as well as between lactating and non lactating females. The reasons for segregation appeared to be predominantly social, but driven ultimately by predation and concomitantly by resources. The habitat selection information was used to design a stratified random sampling strategy that led to i) a significant reduction in survey effort in sampling these sparsely distributed species and ii) reduction in sampling bias.
The applied aspect of the study outlines and evaluates the dramatic changes in the nomadic pastoralism that have occurred in the past five decades in the study area. These have led to a loss of pastures (-25 to -33%) of the nomads, consequent readjustment in traditional patterns of pasture use, intensified grazing pressures (25 to 70%) and rangeland degradation in the area. Such changes may have serious consequences on the survival of local wildlife, as tested with a study of the effects on argali of livestock presence and resource exploitation. Hence, a successful conservation and recovery strategy should focus on: minimising the impacts of livestock on argali, identifying the factors affecting the persistence of the current populations, increasing local sub populations of this species to prevent extinction due to stochastic events, prevent loss of genetic diversity and excessive fragmentation and thus ensuring gene flow.
Ecological Niche Factor Analyses (ENFA), bias-reduced logistic regression and Fuzzy correspondence analyses (FCA) were used to answer habitat and resource selection questions. A sexual segregation and aggregation statistic (SSAS) was used to estimate the components of sexual segregation and test segregation. SSAS combined with canonical correspondence analyses (CCA) allowed the estimation of segregation based on habitat variables. Logistic regression models were formulated to estimate models on which the stratified random sampling strategy was based. The 9 Animal – Habitat relationships in high altitude rangelands overall study also included surveys, interviews and literature reviews to understand the nomads’ movement and pasture use patterns of their livestock. Kernel density estimations (KDE) were used to estimate extent of range overlaps between livestock and argali.
Keywords: high altitude, homogeneous, argali, habitat selection, resource selection function, ENFA, stratified random sampling, sexual segregation, SSAS, livestock, predation, resources
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Ale, S. B. (1997). The Annapurna Conservation Area Project: A Case Study of an Integrated Conservation and Development Project in Nepal. In R. Jackson, & A. Ahmad (Eds.), (pp. 155–169). Lahore, Pakistan: Islt.
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Xinchun, M. (1994). Distribution in the wild and the captive raising of snow leopards in Xinjiang, China. In J.L.Fox, & D.Jizeng (Eds.), (pp. 157–162). Usa: Islt.
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Shi, K., Jun, Z. F. S., Zhigang, D., Riordan, P., & MacDonald, D. (2009). Reconfirmation of snow leopards in Taxkurgan Nature Reserve, Xinjiang, China. Oryx, 43(2), 169–170.
Abstract: China may hold a greater proportion of the global snow leopard Panthera uncia population than any other country, with the area of good quality suitable habitat, estimated at nearly 300,000 km2, comprising .50% of that available across the species' entire range. We can now reconfirm the presence of snow leopard in the Taxkurgan area of Xinjiang Province in north-west China after a period of 20 years.
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Ferguson, D. A. (1997). International Cooperation for Snow Leopard and Biodiversity Conservation: The Government Perspective. In R.Jackson, & A.Ahmad (Eds.), (pp. 178–193). Lahore, Pakistan: Islt.
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Subbotin, A. E., & Istomov, S. V. (2009). The population status of snow leopards Uncia uncia (Felidae, Carnivora) in the western Sayan Mountain Ridge. Doklady Biologicl Sciences, 425, 183–186.
Abstract: The snow leopard (Uncia uncial Schreber, 1776) is the most poorly studied species of the cat family in the world and, in particular, in Russia, where the northern periphery of the species area (no more than 3% of it) is located in the Altai-Hangai-Sayan range [1]. It is generally known that the existing data on the Russian part of the snow leopard population have never been a result of targeted studies; at best, they have been based on recording the traces of the snow leopard vital activity [2]. This is explained by the snow leopard's elusive behavior, inaccessibility of its habitats for humans, and its naturally small total numbers in the entire species area. All published data on the population status of the snow leopard in Russia, from the first descriptions of the species [3-6] to the latest studies [7, 8] are subjective, often speculative, and are not confirmed by
quantitative estimates. It is obvious, however, that every accurate observation of this animal is of particular interest [9]. The purpose of our study was to determine the structure and size of the population group presumably inhabiting the Western Sayan mountain ridge at the northern boundary of the species area
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Zhiryakov V.A. (2002). Ecology and behavior of the Snow leopard in Kazakhstan (Vol. N 1-4.).
Abstract: The data on spreading, numbers and population density of snow leopard in Kazakhstan are given in this article. The total number of the snow leopard in Kazakhstan is evaluated in 100-110 individuals. The everywhere occurred numbers' reduction under the influence of the anthropogenic factors is observed. The snow leopard' inhabitation area varies from 20 to 120 square kilometers depending on its regions. Sex and composition of the population and its aggregative behavior are given. The dynamics of numbers and mortality are estimated.
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Li, J., McCarthy, T. M., Wang, H., Weckworth, B. V., Shaller, G. B., Mishra, C., Lu, Z., Beissinger, S. R. (2016). Climate refugia of snow leopards in High Asia. Biological Conservation, (203), 188–196.
Abstract: Rapidwarming in High Asia is threatening its unique ecosystemand endemic species, especially the endangered
snow leopard (Panthera uncia). Snow leopards inhabit the alpine zone between snow line and tree line, which
contracts and expands greatly during glacier-interglacial cycles. Here we assess impacts of climate change on
global snow leopard habitat from the last glacial maximum (LGM; 21 kyr ago) to the late 21st century. Based
on occurrence records of snow leopards collected across all snow leopard range countries from 1983 to 2015,
we built a snow leopard habitat model using the maximum entropy algorithm (MaxEnt 3.3.3k). Then we
projected this model into LGM, mid-Holocene and 2070. Analysis of snow leopard habitat map from LGM to
2070 indicates that three large patches of stable habitat have persisted from the LGM to present in the Altai,
Qilian, and Tian Shan-Pamir-Hindu Kush-Karakoram mountain ranges, and are projected to persist through the
late 21st century. These climatically suitable areas account for about 35% of the snow leopard's current extent,
are large enough to support viable populations, and should function as refugia for snow leopards to survive
through both cold and warm periods. Existence of these refugia is largely due to the unique mountain environment
in High Asia, which maintains a relatively constant arid or semi-arid climate. However, habitat loss leading
to fragmentation in the Himalaya and Hengduan Mountains, as well as increasing human activities, will present
conservation challenges for snow leopards and other sympatric species.
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Green, M. J. B., & Zhimbiev, B. (1997). Transboundary Protected Areas and Snow Leopard Conservation. In R.Jackson and A.Ahmad (Ed.), (pp. 194–202). Lahore, Pakistan: Islt.
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Mallon, D. P., & Nurbu, C. (1988). A Conservation Program for the Snow Leopard in Kashmir. In H.Freeman (Ed.), (pp. 207–214). India: International Snow Leopard Trust and The Wildlife Institute of India.
Abstract: This program was drawn up at the invitation of the Chief Wildlife Warden of the State of Jammu and Kashmir, Mir Inayat Ullah, following joint survey work and discussions held over the period 1980-86. The recomendations have been accepted and will be incorporated into Department of Wildlife Protection policy. The aim of the program is the long term conservation of the snow leopard on a self sustaining basis throughout the state. Two basic assumptions underlie the program: (1) The snow leopard cannot be protected in isolation, but only in the context of conservation of the environment as a whole, and (2) To be most effective, any long term plan needs the cooperation of local people and must take into account their needs and traditional rights.
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Xu, F., Ming, M., Yin, S. -jing, & Munkhtsog, B. (2006). Autumn Habitat Selection by Snow Leopard (Uncia uncia) in Beita Mountain, Xinjiang, China.
Abstract: Habitat selection of Snow Leopard ( Unica unica) in Beita Mountain of the Altay Mountain system in northeast Xinjiang was conducted from September to October 2004. Six habitat features of 59 sites used by Snow Leopard and 30 random plots were measured by locating 15 transects surveys in the study area . Vanderploge and Scaviaps selectivity index was used to assess Snow Leopardps selection for the different habitat parameters. Principal Component Analysis was used as the primary factor . The results indicated that Snow Leopard preferred the altitude between 2000 – 2 200 m and avoided 2 600 – 3 000 m ; selected cliff base , ridgeline and avoided hillside and valley bottom ; utilized the shrub and rejected the forest ; selected the nongrazing area and avoided the slightly broken region ; preferred north orientation and rejected the south orientation. The results show that grazing status , vegetation type , topography and the ruggedness are the primary factors for the habitat selection of Snow Leopard.
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Green, M. J. B. (1994). Protecting the mountains of Central Asia and their snow leopard populations. In J.L.Fox, & Du Jizeng (Eds.), (pp. 223–239). International Snow Leopard Trust and Chicago Zoological Society.
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Braden, K. E. (1988). Economic Development in Six Regions of Snow Leopard Habitat in the U.S.S.R. In H.Freeman (Ed.), (pp. 227–246). India: International Snow Leopard Trust and the Wildlife Institute of India.
Abstract: The Disappearance of traditional ungulate prey of the snow leopard may be contributing to its endangered status in the wild. Soviet biologists have noted that wild sheep are a primary prey of the snow leopard in the southern Russian union republic and the Central Asian union republic of the U.S.S.R. While poaching appears to have had some impact on the status of these sheep, econmic pressures may be contributing to their decrease. Evidence presented for KAzakhstan and three regions of the Russian republic demonstrates that commercial sheep and goat production appears to be growing at a very high pace in these areas, thus consumming habitat otherwise available for wild herds.
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Jackson, R. M. (1996). Home Range, Movements and Habitat use of Snow Leopard (Uncia uncia) in Nepal. Ph.D. thesis, University of London, University of London.
Abstract: Home ranges for five radio-tagged snow leopards (Uncia uncia) inhabiting prime habitat in Nepal Himalaya varied in size from 11-37 km2. These solitary felids were crepuscular in activity, and although highly mobile, nearly 90% of all consecutive day movements involved a straight line distance of 2km or less. No seasonal difference in daily movement or home range boundry was detected. While home ranges overlapped substancially, use of common core spaces was temporally seperated, with tagged animals being located 1.9 km or more apart during the smae day. Spatial analysis indicated that 47-55% of use occured within only 6-15% of total home area. The snow leopards shared a common core use area, which was located at a major stream confuence in an area where topography, habitat and prey abundance appeared to be more favorable. A young female used her core area least, a female with two cubs to the greatest extent. the core area was marked significantly more with scrapes, Faeces and other sighn than non-core sites, suggesting that social marking plays an important role in spacing individuals. Snow leopards showed a strong preference for bedding in steep, rocky or broken terrain, on or close to a natural vegetation or landform edge. linear landform features, such as a cliff or major ridgeline, were preferred for travelling and day time resting. This behavior would tend to place a snow leopard close to its preferred prey, blue sheep (Psuedois nayaur), which uses the same habitat at night. Marking was concetrated along commonly travelled routes, particularly river bluffs, cliff ledges and well defined ridgelines bordering stream confluences--features that were most abundant within the core area. Such marking may facilitate mutual avoidance, help maintain the species' solitary social structure, and also enable a relatively high density of snow leopard, especially within high-quality habitat.
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Sokolov V.E. (1979). Snow leopard genus.
Abstract: Genus Uncia has only one species snow leopard. Described is the length of body, length of tail, body weight, colour, skull, number of teeth, habitat, reproduction, and life expectancy. This species has no essential practical value and is included in the Red Data Book as an endangered species.
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Freeman, H., Jackson, R., Hillard, R., & Hunter, D. O. (1994). Project Snow Leopard: a multinational program spearheaded by the International Snow Leopard Trust. In J.L.Fox, & D. Jizeng (Eds.), (pp. 241–245). Usa: Islt.
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Hunter, D. O., Jackson, R., Freeman, H., & Hillard, D. (1994). Project snow leopard: a model for conserving central Asia biodiversity. In J.Fox, & D.Jizeng (Eds.), (pp. 247–252). Usa: International Snow Leopard Trust.
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Villarrubia, C., & Jackson, R. (1994). Snow Leopard Conservation on a Regional Basis: Elements in Planning Protected Areas. In J.L.Fox, & D.Jizeng (Eds.), (pp. 253–263). Usa: Islt.
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Taber, R. (1988). Long Term Research in Snow Leopard Conservation. In H.Freeman (Ed.), (pp. 255–259). India: International Snow Leopard Trust and Wildlife Institute of India.
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Freeman, H. (1988). Resolutions Conservation of Snow Leopard, Fifth International Snow Leopard Symposium. (pp. 267–269). Usa.
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Novikov G.A. (1956). Sub-genus UNCIA Gray.
Abstract: Identification features of the sub-genus Uncia (colour; length of body and tail; shoulder height, and skull measurements) are given. Distribution, habitat, way of life, reproduction biology, behavioural patterns, migration routes, commercial value of snow leopard in the USSR is described.
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Yachontov A.A. (1970). The cats – Felidae.
Abstract: It describes Felidae family species (lion, tiger, leopard, snow leopard, lynx, cheetah): identification features, life-history characteristics, type of habitat, diet and distribution.
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Norberg-Hodge, H. (1981). Ladakh: Developement without Destruction. In J. S. Lall (Ed.), The Himalaya: Aspects of Change (pp. 278–284). New Delhi: Oxford University Press.
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Baryshnikov G.F. (1981). Sub-genus Panthera Oken, 1816. Genus Panthera.
Abstract: The monograph provides taxonomic description of sub-genus Panthera Oken, 1816, genus Uncia grey, 1854. Snow leopard inhabits the mountains of Tajikistan, the Pamirs, Tien Shan, Tarbagatai, the Altai, the Sayans; also the mountain of Mongolia, Tibet, the Himalayas, and Hindukush, where it keeps to alpine meadows and woodless rocks at up to 3,000 – 4,000 m above sea level in summer, and descends to a lower elevation in winter. It described from the Altai. They are of minor trade importance. This species is rare all over its habitat and included in the Red Data Book of the USSR.
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Maheshwari, A., Sharma, D., Sathyakumar, S. (2013). Snow Leopard (Panthera Uncia) surveys in the Western Himalayas, India. Journal of Ecology and Natural Environmnet, 5(10), 303–309.
Abstract: We conducted surveys above 3000 m elevation in eight protected areas of Uttarakhand and Himachal Pradesh. These surveys provide new information on snow leopard in Uttarakhand on the basis of indirect evidence such as pugmark and scat. Snow leopard evidence (n = 13) were found between 3190 and 4115 m elevation. On an average, scats (n = 09) of snow leopard were found for every 56 km walked and pugmarks (n = 04) for every 126 km walked. Altogether, about 39% of the evidence were found on the hill-slope followed by valley floor (30%), cliff (15%) and 8% from both stream bed and scree slope. Genetic analysis of the scats identified three different individuals by using snow leopard specific primers. Snow leopard-human conflicts were assessed through questionnaire based interviews of shepherds from Govind Pashu Vihar Wildlife Sanctuary, Askot Wildlife Sanctuary and Nanda Devi Biosphere Reserve areas of Uttarakhand. Surveys revealed that livestock depredation (mule, goat and sheep) is the only cause of snow leopard-human conflicts and contributed 36% of the diet of snow leopard. Blue sheep and rodents together comprised 36.4% of the total diet. We found that 68.1% of the surveyed area was used for pastoral activities in Uttarakhand and Himachal Pradesh and 12.3% area was under tourism, defence and developmental activities.
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