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Ahmad, S., Rehman, E. U., Ali, H., Din, N., Haider, J., Din, J. U., Nawaz, M. A. (2022). Density Pattern of Flare-Horned Markhor (Capra falconeri) in Northern Pakistan. Sustainability, 14(9567), 1–13.
Abstract: Wild ungulates play vital roles in maintaining a balanced ecosystem through herbivory and are also an important determinant of carnivores’ density. The flare-horned markhor (Capra falconeri) is a threatened wild goat distributed across the mountain ranges of Pakistan, India, Afghanistan, Russia, Turkmenistan, Uzbekistan, and Tajikistan. The remote terrain and fragmented population limit our understanding of the population ecology of markhor, though knowledge of the target species population is vital for making informed management decisions. Therefore, the current study was designed to determine the markhor population across their range in Northern Pakistan and to evaluate the efforts made by the government and non-government organizations for the conservation of markhor. Double-observer surveys were conducted during 2019–2021 in nine major watersheds of Khyber Pakhtunkhwa and Gilgit-Baltistan covering an area of 4664 km2. Secondary data were collected for unassessed areas to gain a holistic overview of the markhor population and density in the region. Results revealed a markhor population of 7579, with a density of 0.30 animals per km2 in Northern Pakistan. Our analysis of the double-observer data through the Bayesian behavioral capture–recapture model estimated a population of 5993 individuals (95% CI) of markhor across
nine study sites, with a density of 1.28 animals per km . A review of secondary data revealed that a population of about 1586 was present in the un-surveyed area (20,033.33 km2), with a density of 0.08 per km . A total of 146 groups of markhor were counted, with a mean group size of 23 (3–58) individuals. There were 109 males and 108 young per 100 females in the population. Among 1936 recorded males, Class I males accounted for 27.74%, followed by Class II (26.45%), Class IV (trophy-size) (23.40%), and Class III (22.42%). The overall detection probability was recorded as 0.87 and 0.68 for the first observer and second observer, respectively. Compared with the past reports, the population of markhor in Northern Pakistan appears to be increasing, particularly in protected areas (PAs) such as national parks and community-controlled hunting areas (CCHAs). Conservation programs, notably trophy hunting and PA networks, appear to be vital in sustaining markhor populations in parts of the species range. We recommend expansion in such programs in the markhor range in order to maintain a viable population of this majestic wild goat in the region. Keywords: markhor; Capra falconeri; Gilgit-Baltistan; Karakoram; population; double-observer; CGNP
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Ahmad, S., Ali, H., Asif, M., Khan, T, Din, N., Rehman, E. U., Hameed, S., Din, J. U., Nawaz, M. A. (2022). Spatial density pattern of Himalayan Ibex (Capra sibirica) in Pakistan. Global Ecology & Conservation, 39(e02288), 1–12.
Abstract: Mountain ungulates perform a key role in maintaining the balance of ecosystems as they are the primary consumers of vegetation and prey for large predators. The mountain ranges of northern Pakistan are home to six species of mountain ungulates, and the Himalayan ibex (Capra sibirica), hereafter ibex, is the most abundant among them. This study was conducted in three administrative regions of northern Pakistan, viz. Gilgit-Baltistan (GB), Azad Jammu and Kashmir (AJK), and Khyber Pakhtunkhwa (KP), to generate a range-wide density pattern map of ibex. A double-observer survey was conducted in 25 study sites during 2018–2021 across the ibex distribution range, covering an area of about 35,307 km2, by walking transects totaling 1647 km. Within the ibex range where the survey was not conducted due to financial and logistical constraints, we obtained species population information from local wildlife departments’ most recent annual survey data. The aim was to generate a density map for the entire ibex range. Using the BBRe-capture package in program R, we estimated an ibex population of 7639 (95 % CI) with a mean density of 0.21/km2 in the surveyed area. Combining with the secondary data from un-surveyed areas, the total population estimate for the country came to 10,242 ibex. The largest population densities were observed in four valleys (Shimshal, Gulkin-Hussaini, Khyber, and Khunjerab) of the Karakoram-Pamir range, followed by the Hindu Kush range (Chitral Wildlife Division [WD]). The central and eastern parts of the Karakoram range had moderate to low densities, while the Himalayan range (e.g., Astore Valley) supported a small population. The mean herd size was 15 individuals (range: 5–41), and the average detection probability of observers A and B was 0.69 and 0.48, respectively. The average male and young ratios per 100 females were estimated to be 75 and 81, respectively. The range-wide density map developed during the study provided an evidence for the impact of trophy hunting programs and an objective tool for range-wide conservation planning of the species.
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Ale S. (2005). Have snow leopards made a comeback to the Everest region of Nepal?.
Abstract: In the 1960s, the endangered snow leopard was locally extirpated from the Sagarmatha (Mt. Everest) region of Nepal. In this Sherpa-inhabited high Himalaya, the flourishing tourism since the ascent of Mt Everest in 1953, has caused both prosperity and adverse impacts, the concern that catalyzed the establishment of Mt. Everest National Park in the region in 1976. In the late 1980s, there were reports that some transient snow leopards may have visited the area from adjoining Tibet, but no biological surveys exist to confirm the status of the cats and their prey. Have snow leopards finally returned to the top of the world? Exploring this question was the main purpose of this research project. We systematically walked altogether 24 sign transects covering over 13 km in length in three valleys, i.e. Namche, Phortse and Gokyo, of the park, and counted several snow leopard signs. The results indicated that snow leopards have made a comeback in the park in response to decades of protective measures, the virtual cessation of hunting and the recovery of the Himalayan tahr which is snow leopard's prey. The average sign density (4.2 signs/km and 2.5 sign sites/km) was comparable to that reported from other parts of the cats' range in the Himalaya. On this basis, we estimated the cat density in the Everest region between 1 to 3 cats per 100 sq km, a figure that was supported by different sets of pugmarks and actual sightings of snow leopards in the 60 km2 sample survey area. In the study area, tahr population had a low reproductive rate (e.g. kids-to-females ratio, 0.1, in Namche). Since predators can influence the size and the structure of prey species populations through mortality and through non-lethal effects or predation risk, snow leopards could have been the cause of the population dynamics of tahr in Sagarmtha, but this study could not confirm this speculation for which further probing may be required.
Keywords: snow; snow leopards; snow leopard; snow-leopards; snow-leopard; leopards; leopard; region; Nepal; Report; International; international snow leopard trust; International-Snow-Leopard-Trust; trust; program; 1960; endangered; Sagarmatha; High; Himalaya; tourism; impact; establishment; national; national park; National-park; park; 1980; area; Tibet; surveys; survey; status; Cats; cat; prey; research; project; sign; transects; transect; length; valley; Response; hunting; recovery; Himalayan; tahr; density; densities; range; pugmarks; sighting; 60; study; population; predators; predator; structure; prey species; prey-species; species; populations; mortality; effects; predation; population dynamics
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Jackson, R. (2002). Snow Leopard Status, Distribution, and Protected Areas Coverage.. Islt: Islt.
Abstract: This document reports on the status, distribution and protected areas coverage for snow leopard across its range in Central Asia. It is intended to aid in updating the existing knowledge base of snow leopard status and distribution during the Snow Leopard Survival Strategy (SLSS) Workshop organized by the International Snow Leopard Trust (ISLT), and to be held in Seattle on May 21-25, 2002. The SLSS workshop provides an unique opportunity to solicit feedback from scientists,
conservation organizations, government agencies and knowledgeable experts in order to: (1) Verify the accuracy of information presented in this document (much of which was published 5-10 years ago) and to identify data gaps, especially with respect to population size and protectedareas coverage for this species; (2) Aid organizations in developing tightly targeting conservation actions by identifying critical snow leopard areas, parks and reserves, and by implication, the intervening linking corridors linking key protected areas; (3) Serve as a catalyst for encouraging range-countries to conduct field surveys into snow leopard status and distribution, especially in those areas deemed the most important to the maintenance of a viable metapopulation across the 12 countries in which the species' occurs. Keywords: snow; leopard; status; distribution; range; China; Bhutan; conservation; protection; protected; protect; areas; Afghanistan; Nepal; Ussr; population; kyrgystan; Pakistan; Uzbekistan; Kazakhstan; Tajikistan; 4980
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Allen, P., & Macray, D. (2002). Snow Leopard Enterprises Description and Summarized Business Plan.. Seattle: Islt.
Abstract: The habitat for both humans and snow leopards in Central Asia is marginal, the ecosystem fragile. The struggle for humans to survive has often, unfortunately, brought them into conflict with the region's dwindling snow leopard populations. Herders commonly see leopards as a threat to their way of life and well-being. Efforts to improve the living conditions of humans must consider potential impacts on the environment. Likewise, conservation initiatives cannot ignore humans as elements of the landscape with a right to live with dignity and pride. Based on these principles, the International Snow Leopard Trust has developed a new conservation model that addresses the needs of all concerned.
We call it Snow Leopard Enterprises.. Keywords: snow; leopard; enterprises; buisness; plan; habitat; humans; conflict; irbis; products; wool; conservation; marketing; Mongolia; social; economic; conflicts; country; countries; socks; hats; gloves; 4890; Human; snow leopards; snow leopard; snow-leopards; snow-leopard; leopards; central; Central Asia; asia; ecosystem; region; populations; population; herders; herder; threat; potential; impact; environment; Elements; landscape; International; international snow leopard trust; International-Snow-Leopard-Trust; trust; snow-leopard-enterprises
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Anonymous. (1999). Livestock Predation Control Workshop.
Keywords: Lahul-Spiti; Ladakh; Hemis; parks; reserves; refuge; protected-area; argali; abix; blue-sheep; wolves; distribution; status; population; poaching; hunting; trade; skins; livestock; pelts; coat; fur; bones; medicine; prey-depletion; herders; habitat; habitat-degradation; tourism; Tmi; Islt; predator; prey; conflict; compensation; trekking; blue; sheep; browse; protected; area; depletion; degradation; international snow leopard trust; 3940
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Aripjanov M.P. (1990). Rare mammals of South-West Tien Shan.
Abstract: Rare mammal species such as free-toiled bat, Menzbier's marmot (endemic to the Western Tien Shan), Tien-Shan brown bear, Central Asian otter, Turkestan lynx, snow leopard, and wild sheep inhabit the South-West Tien-Shan (Uzbekistan). Brief data on animal encounters and main threats are given.
Keywords: Uzbekistan; South-West Tien-Shan; rare species; snow leopard; population; species number; poaching; human activity.; 6040; Russian
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Aromov, B. (2001). Snow Leopard (Uncia uncia) in Hissar Nature Reserve (Vol. Issue 3).
Abstract: Data on distribution, number, diet and breeding of snow leopard in NW spurs of the Hissar Ridge were collected over long-term studies in the span from 1981 to 1994. An increase in the number of this animal from 4 to 17 individuals has been recorded in the Hissar Nature Reserve (Uzbekistan).
Keywords: Uzbekistan; snow leopard; Hissar ridge; Hissar nature reserve; number; population estimate; diet; wild ibex; livestock; rut; cubs; competitors.; 6080; Russian
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Blomqvist, L. (1982). The 1981 annual report of the captive snow leopards (Panthera uncia) population. International Pedigree Book of Snow Leopards, 3. |
Chapron, G., & Legendre, S. (2002). Some Insights Into Snow Leopard (Uncia Uncia) Demography By Using Stage Structured Population Models.. Seattle: Islt.
Abstract: Based on the limited data available on snow leopard demography, we developed deterministic and stochastic stage-structured demographic models to study the population dynamics of this large cat. Our results reveal that even small leopard populations can persist provided their demographic parameters remain high, but less favorable scenarios would require larger population sizes. Population growth rate is more sensitive to breeder survivals than to any other parameters. A snow leopard population would start declining if yearly mortality claims more than 1/5 of the population. This study identifies poaching as a major threat to snow leopard survival and stresses the importance of long-term studies to better understand snow leopard population dynamics.
Keywords: snow; leopard; uncia; viability; analysis; carnivore; carnivores; domestic; game; demographic; population; mortality; biology; mating; 4910
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Fox, J. L., & Jackson, R. M. (2002). Blue Sheep and Snow Leopards in Bhutan and Trans-Himalayan Nepal: Recent Status Evaluations and Their Application to Research and Conservation.. Islt: Islt. |
Jackson, R., & Ahlborn, G. (1990). The role of protected areas in Nepal in maintaining viable populations of snow leopards. Int.Ped.Book of Snow Leopards, 6, 51–69. |
Jackson, R., & Fox, J. L. (2000). Report on Fifth Slims Training Workshop (Nepal) (Vol. xvii). Seattle: International Snow Leopard Trust.
Abstract: Nepal's snow leopards (Uncia uncia) are mostly found along the northern border with Tibet (China). The largest populations are in Dolpa, Mugu, Manang, and Myagdi Districts. Potential habitat totals about 30,000 square kilometers. Numbers are estimated at 300-500, but surveys are urgently needed to confirm this rough guess. Like elsewhere, the primary threats center on poaching, depletion of natural prey, livestock depredation and resultant retributive killing of snow leopards by herders, and the lack of public awareness and support for conserving snow leoaprds, especially among local herders.
Keywords: Slims; Nepal; training; techniques; Gps; field-work; surveys; Tibet; habitat; China; hunting; poaching; livestock; population; Shey-Phoksundo; parks; protected-area; reserves; annapurna; Dhorpatan; Manaslu; Sagarmatha; Langtang; Islt; Wwf; Hmg; Dnpwc; browse; 4460
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Jackson, R., & Roe, J. (2002). Preliminary Observations On Non-Invasive Techniques for Identifying Individual Snow Leopards and Monitoring Populations.. Islt: Islt. |
Sokov, A. I. (1990). The present status of the snow leopard population in the south western Pamir-Altai Mountains (Tadzhikistan). Int.Ped.Book of Snow Leopards, 6, 33–36. |
Jackson, R., Roe, J., Wangchuk, R., & Hunter, D. (2005). Surveying Snow Leopard Populations with Emphasis on Camera Trapping: A Handbook. Sonoma, California: The Snow Leopard Conservancy.
Abstract: This handbook provides an introduction to snow leopard population survey techniques, followed by a detailed account of camera trapping methods.During the 2002 through 2004 winter field seasons, the Snow Leopard Conservancy experimented with infrared camera trapping techniques to define a methodology suitable for the high altitude environment.
In 2001 and 2002, much of our time was spent familiarizing ourselves with various infrared camera traps, their operation and setup, and comparing the effectiveness of different models and sensor types. We placed infrared camera traps along frequently used travel corridors at or near scent-sprayed rocks (rock scents) and scrape sites within 16 km2 sampling cells between January and March in 2003 and 2004. A total of 66 and 49 captures of snow leopards were tallied during 2003 and 2004, resulting in an overall capture success of 8.91 and 5.63 individuals per 100 trap-nights, respectively. Capture probabilities ranged from 0.33 to 0.46. Density estimates ranged from 8.49 ± 0.22 individuals per 100 km2 in 2003 to 4.45 ± 0.16 in 2004, with the disparity between years largely attributed to different trapping densities. Snow leopard abundance estimates were calculated using the computer program CAPTURE. Keywords: snow; snow leopard; snow-leopard; leopard; survey; conservation; populations; population; camera; camera trapping; trapping; Chinese
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Jackson, R., Roe, J., Wangchuk, R., & Hunter, D. (2005). Surveying Snow Leopard Populations with Emphasis on Camera Trapping: A Handbook. Sonoma, California: The Snow Leopard Conservancy.
Abstract: This handbook provides an introduction to snow leopard population survey techniques, followed by a detailed account of camera trapping methods.During the 2002 through 2004 winter field seasons, the Snow Leopard Conservancy experimented with infrared camera trapping techniques to define a methodology suitable for the high altitude environment.
In 2001 and 2002, much of our time was spent familiarizing ourselves with various infrared camera traps, their operation and setup, and comparing the effectiveness of different models and sensor types. We placed infrared camera traps along frequently used travel corridors at or near scent-sprayed rocks (rock scents) and scrape sites within 16 km2 sampling cells between January and March in 2003 and 2004. A total of 66 and 49 captures of snow leopards were tallied during 2003 and 2004, resulting in an overall capture success of 8.91 and 5.63 individuals per 100 trap-nights, respectively. Capture probabilities ranged from 0.33 to 0.46. Density estimates ranged from 8.49 ± 0.22 individuals per 100 km2 in 2003 to 4.45 ± 0.16 in 2004, with the disparity between years largely attributed to different trapping densities. Snow leopard abundance estimates were calculated using the computer program CAPTURE. Keywords: snow; snow leopard; snow-leopard; leopard; populations; population; camera; camera trapping; trapping
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Mongolian Biosphere & Ecology Association. (2010). Mongolian Biosphere & Ecology Association Report March 2010.
Abstract: In accordance with order of the Ministry of Nature and Tourism,
zoologists of our association have made surveys in three ways such as reasons why snow leopards attack domestic animals, “Snow leopard” trial operation to count them and illegal hunting in territories of Khovd, Gobi-Altai, Bayankhongor, Uvurkhangai and Umnugobi provinces from September 2009 to January 2010. As result of these surveys it has made the following conclusions in the followings: Reason to hunt them illegally: the principal reason is that administrative units have been increased and territories of administrative units have been diminished. There have been four provinces in 1924 to 1926, 18 since 1965, 21 since 1990. Such situation limits movements of herdsmen completely and pastures digressed much than ever before. As result of such situation, 70% of pastures become desert. Such digression caused not only heads of animals and also number of species. Guarantee is that birds such as owls, cuckoo, willow grouse in banks of Uyert river, Burkhanbuudai mountain, located in Biger soum, Gobi-Altai province, which are not hunted by hunters, are disappearing in the recent two decades. For that reason we consider it is urgently necessary for the government to convert administrative unit structures into four provinces. This would influence herdsmen moving across hundreds km and pastures could depart from digression. Second reason: cooperative movement won. The issues related to management and strengthening of national cooperatives, considered by Central Committee of Mongolian People's Revolutionary Party in the meeting in March 1953 was the start of cooperatives' movement. Consideration by Yu. Tsedenbal, chairman of Ministers Council, chairman of the MPRP, on report "Result of to unify popular units and some important issues to maintain entity management of agricultural cooperatives" in the fourth meeting by the Central Committee of Mongolian People's Revolutionary Party /MPRP/ on December 16-17, 1959, proclaimed complete victory of cooperative. At the end of 1959, it could unify 767 small cooperative into 389 ones, unify 99.3 % of herdsmen and socialize 73.3 % of animals. The remaining of animals amount 6 million 163 thousands animals, and equals to 26.7% of total animals. This concerned number of animals related to the article mentioned that every family should have not more that 50 animals in Khangai zone and not more 75 animals in Gobi desert. It shows that such number could not satisfy needs of family if such number is divided into five main animals in separating with reproduction animals and adult animals. So herdsmen started hunt hoofed animals secretly and illegally in order to satisfy their meat needs. Those animals included main food of snow leopard such as ibex, wild sheep, and marmot. Third reason is that the state used to hunt ibex, which are main nutrition of snow leopards, every year. The administrative unit of the soum pursued policy to hunt ibex in order to provide meat needs of secondary schools and hospitals. That's why this affected decrease of ibex population. Preciously from 1986 to 1990 the permissions to hunt one thousands of wild sheep and two thousands of ibexes were hunt for domestic alimentary use every year. Not less than 10 local hunters of every soum used to take part in big game of ibexes. Also they hunted many ibexes, chose 3-10 best ibexes and hid them in the mountains for their consummation during hunting. Fourth reason: hunting of wolves. Until 1990 the state used to give prizes to hunter, who killed a wolf in any seasons of the year. Firstly it offered a sheep for the wolf hunter and later it gave 25 tugrugs /15 USD/. Every year, wolf hunting was organized several times especially picking wolf-cubs influenced spread and population of wolves. So snow leopard came to the places where wolves survived before and attack domestic animals. Such situation continued until 1990. Now population of ibexes has decreased than before 1990 since the state stopped hunting wolves, population of wolves increased in mountainous zones. We didn't consider it had been right since it was natural event. However population of ibexes decreased. Fifth reason: Global warming. In recent five years it has had a drought and natural disaster from excessive snow in the places where it has never had such natural disasters before. But Mongolia has 40 million heads of domestic animals it has never increased like such quantity in its history before. We consider it is not incorrect that decrease of domestic animals could give opportunities to raise population of wild animals. Our next survey is to make attempt to fix heads of snow leopards correctly with low costs. Keywords: nature; tourism; surveys; survey; snow; snow leopards; snow leopard; snow-leopards; snow-leopard; leopards; leopard; attack; domestic; Animals; Animal; illegal; illegal hunting; hunting; territory; province; 2010; hunt; 1990; movements; movement; pasture; desert; number; species; birds; river; mountain; hunters; hunter; recent; government; structure; management; national; central; people; Report; gobi; Gobi Desert; reproduction; Adult; meat; food; ibex; wild; wild sheep; sheep; marmot; nutrition; schools; population; use; local; big; big game; big-game; game; 310; mountains; wolves; wolf; Seasons; times; zones; global; Mongolia; 40; history; ecology
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Smirnov, M. N., Sokolov, G. A., & Zyryanov, A. N. (1990). The Snow Leopard (Uncia Uncia Scherber 1776) in Siberia. Int.Nat.Ped.Book of Snow Leopards, 6, 9–15.
Keywords: siberia; status; distribution; Russia; Ussr; Soviet-Union; browse; soviet union; soviet; union; Altai; Sayan; population; tracks; tracking; petroglyphs; skins; pelts; prey; 2880
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Saltz, D., Rowen, M., & Rubenstein, D. (2000). The effect of space-use patterns of reintroduced Asiatic wild ass on effective population size. Conservation Biology, 14(6), 1852–1861. |
Reading, R. P., Amgalanbaatar, S., Mix, H., & Lhagvasuren, B. (1997). Argali Ovis ammon surveys in Mongolia's South Gobi. Oryx, 31(4), 285–294.
Abstract: Claims poaching and competition with domestic livestock are threatening the argali's survival in Mongolia. The author's conducted aerial and ground surveys in the South Gobi and estimated a populaton size of approximately 3,900 argali.
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Poyarkov, A. D., & Subbotin, A. E. (2002). Strategic Priorities and the System of Measures for Snow Leopard Conservation in Russia.. Islt: Islt. |
Poyarkov, A. D., & Subbotin, A. E. (2002). The Snow Leopard Status in Russia.. Islt: Islt.
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Poyarkov, A. D. (2002). Some Aspects of Snow Leopard Research Methodology.. Islt: Islt. |
Koshkarev, E. (2002). Strategy of Snow Leopard Conservation in the Range.. Islt: Islt. |