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Author Namgail, T.; Fox, J.; Bhatnagar, Y.V. url 
  Title Habitat segregation between sympatric Tibetan argali Ovis ammon hodgsoni and blue sheep Pseudois nayaur in the Indian Trans-Himalaya Type Journal Article
  Year 2004 Publication Journal of Zoology Abbreviated Journal  
  Volume 262 Issue Pages 57-63  
  Keywords argali; Ovis ammon hodgsoni; blue sheep; pseudois nayaur; Habitat selection; resource partitioning; niche relationship; 5200  
  Abstract Tibetan argali Ovis ammon hodgsoni and blue sheep Pseudois nayaur have almost completely overlapping distributions encompassing most of the Tibetan plateau and its margins. Such a sympatric distribution of related species with similar ecological requirements implies that there is some degree of resource partitioning. This may be accomplished on the basis of habitat and/or diet separation. This study evaluated such ecological separation on the basis of physical habitat partitioning by these two sympatric ungulates in Hemis High Altitude National Park, Ladakh, India, in an area where the argali established a small new population in 1978. Such separation was tested for

on the basis of expected difference between the species in their proximity to cliffs, associated with species-specific anti-predator behaviour. Tibetan argali selected habitats away from cliffs while blue sheep selected habitats close to cliffs. Blue sheep also selected steep slopes whereas argali selected gentle slopes. The two species did not differ

in their use of habitats in terms of elevation. They did, however, differ in their use of plant communities; blue sheep selected sub-shrub and grass-dominated communities whilst argali selected forb-dominated communities. We suggest that the two species coexist in this site as a result of the differential use of habitat associated with their

species-specific anti-predator strategies.
 
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  Notes Full text available at URL Approved no  
  Call Number SLN @ rana @ 505 Serial 710  
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Author Oli, M. url 
  Title Snow leopards and blue sheep in Nepal: Densities and predator: Prey ratio Type Miscellaneous
  Year 1994 Publication Journal of Mammalogy Abbreviated Journal  
  Volume 75 Issue Pages 998-1004  
  Keywords snow leopard,Panthera uncia,blue sheep,Pseudois nayaur,density,predator:prey ratio,harvest rate,livestock predation,Nepal  
  Abstract I studied snow leopards (Panthera uncia) and blue sheep (Pseudois nayaur) in Manang District, Annapurna Conservation Area, Nepal, to estimate numbers and analyze predatorprey interactions. Five to seven adult leopards used the 105-km2 study area, a density of 4.8 to 6.7 leopards/100 km2. Density of blue sheep was 6.6-10.2 sheep/km2, and biomass density was 304 kg/km2. Estimated relative biomass consumed by snow leopards suggested that blue sheep were the most important prey; marmots (Marmota himalayana) also contributed significantly to the diet of snow leopards. Snow leopards in Manang were estimated to harvest 9-20% of total biomass and 11-24% of total number of blue sheep annually. Snow leopard :blue sheep ratio was 1 :1 14-1 :159 on a weight basis, which was considered sustainable given the importance of small mammals in the leopard's diet and the absence of other competing predators.  
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  Notes Approved no  
  Call Number SLN @ rana @ 894 Serial 741  
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Author Oli, M.K. url 
  Title Snow leopards and blue sheep in Nepal: Densities and predator: prey ratio Type Journal Article
  Year 1994 Publication Journal of Mammalogy Abbreviated Journal  
  Volume 75 Issue 4 Pages 998-1004  
  Keywords Nepal; blue-sheep; prey; livestock; predation; blue; sheep; browse; 740; snow; snow leopards; snow leopard; snow-leopards; snow-leopard; leopards; leopard; blue sheep; densities; density; predator  
  Abstract I studied snow leopards (Panthera uncia) and blue sheep (Pseudois nayaur) in Manang District, Annapurna Conservation Area, Nepal, to estimate numbers and analyze predator-prey interactions. Five to seven adult leopards used the 10-5-km-2 study area, a density of 4.8 to 6.7 leopards/100 km-2. Density of blue sheep was 6.6 10.2 sheep/km-2, and biomass density was 304 kg/km-2. Estimated relative biomass consumed by snow leopards suggested that blue sheep were the most important prey; marmots (Marmota himalayana) also contributed significantly to the diel of snow leopards Snow leopards in Manang were estimated to harvest 9-20% of total biomass and 11-24% of total number of blue sheep annually. Snow leopard: blue sheep ratio was 1:114-1:159 on a weight basis, which was considered sustainable given the importance of small mammals in the leopard's diet and the absence of other competing predators.  
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  Notes Document Type: English Call Number: 599.05 JO Approved no  
  Call Number SLN @ rana @ 236 Serial 746  
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Author Oli, M.K.; Taylor, K.R.; Rogers, M.E. url 
  Title Snow leopard Panthera uncia predation of livestock: An assessment of local perceptions in the Annapurna Conservation Area, Nepal Type Miscellaneous
  Year 1994 Publication Biological Conservation Abbreviated Journal  
  Volume 68 Issue Pages 63-68  
  Keywords snow leopard,blue sheep,livestock predation,public attitudes,Annapurna Conservation Area,Nepal.  
  Abstract Public attitudes towards snow leopard Panthera uncia predation of domestic livestock were investigated by a questionnaire survey of four villages in snow leopard habitat within the Annapurna Conservation Area, Nepal Most local inhabitants were subsistence farmers, many dependent upon yaks, oxen, horses and goats, with an average livestock holding of 26.6 animals per household. Reported losses to snow leopards averaged 0.6 and O. 7 animals per household in two years of study, constituting 2.6% of total stockholding but representing in monetary terms almost a quarter of the average annual Nepali national per capita income. Local people hem strongly negative attitudes towards snow leopards and most suggested that total extermination of leopards was the only acceptable solution to the predation problem. Snow leopards were reported to be killed by herdsmen in defence of their livestock. The long-term success of snow leopard conservation programmes may depend upon the satisfactory resolution of the predation conflict. Some possible ways of reducing predation losses are also discussed.  
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  Notes Approved no  
  Call Number SLN @ rana @ 893 Serial 753  
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Author Shrestha, B. url 
  Title Prey Abundance and Prey Selection by Snow Leopard (uncia uncia) in the Sagarmatha (Mt. Everest) National Park, Nepal Type Report
  Year 2008 Publication Abbreviated Journal  
  Volume Issue Pages 1-35  
  Keywords project; snow; snow leopard; snow-leopard; leopard; network; conservation; program; prey; abundance; selection; uncia; Uncia uncia; Uncia-uncia; Sagarmatha; national; national park; National-park; park; Nepal; resource; predators; predator; ecological; impact; region; community; structure; number; research; population; status; density; densities; wild; prey species; prey-species; species; Himalayan; tahr; musk; musk-deer; deer; game; birds; diet; livestock; livestock depredation; livestock-depredation; depredation; awareness; co-existence; ungulates; ungulate; Human; using; areas; area; monitoring; transect; Hair; identification; scat; attack; patterns; sighting; 1760; populations; birth; Male; Female; young; domestic; domestic livestock; 120; scats; yak; Dog; pika; wildlife; Seasons; winter; horse; study; cover; land; predation; Pressure; development; strategy; threatened; threatened species; threatened-species; conflicts; conflict; people; control; husbandry; compensation; reintroduction; blue; blue sheep; blue-sheep; sheep; free ranging  
  Abstract Predators have significant ecological impacts on the region's prey-predator dynamic and community structure through their numbers and prey selection. During April-December 2007, I conducted a research in Sagarmatha (Mt. Everest) National Park (SNP) to: i) explore population status and density of wild prey species; Himalayan tahr, musk deer and game birds, ii) investigate diet of the snow leopard and to estimate prey selection by snow leopard, iii) identify the pattern of livestock depredation by snow leopard, its mitigation, and raise awareness through outreach program, and identify the challenge and opportunities on conservation snow leopard and its co-existence with wild ungulates and the human using the areas of the SNP. Methodology of my research included vantage points and regular monitoring from trails for Himalayan tahr, fixed line transect with belt drive method for musk deer and game birds, and microscopic hair identification in snow leopard's scat to investigate diet of snow leopard and to estimate prey selection. Based on available evidence and witness accounts of snow leopard attack on livestock, the patterns of livestock depredation were assessed. I obtained 201 sighting of Himalayan tahr (1760 individuals) and estimated 293 populations in post-parturient period (April-June), 394 in birth period (July -October) and 195 November- December) in rutting period. In average, ratio of male to females was ranged from 0.34 to 0.79 and ratio of kid to female was 0.21-0.35, and yearling to kid was 0.21- 0.47. The encounter rate for musk deer was 1.06 and density was 17.28/km2. For Himalayan monal, the encounter rate was 2.14 and density was 35.66/km2. I obtained 12 sighting of snow cock comprising 69 individual in Gokyo. The ratio of male to female was 1.18 and young to female was 2.18. Twelve species (8 species of wild and 4 species of domestic livestock) were identified in the 120 snow leopard scats examined. In average, snow leopard predated most frequently on Himalayan tahr and it was detected in 26.5% relative frequency of occurrence while occurred in 36.66% of all scats, then it was followed by musk deer (19.87%), yak (12.65%), cow (12.04%), dog (10.24%), unidentified mammal (3.61%), woolly hare (3.01%), rat sp. (2.4%), unidentified bird sp. (1.8%), pika (1.2%), and shrew (0.6%) (Table 5.8 ). Wild species were present in 58.99% of scats whereas domestic livestock with dog were present in 40.95% of scats. Snow leopard predated most frequently on wildlife species in three seasons; spring (61.62%), autumn (61.11%) and winter (65.51%), and most frequently on domestic species including dog in summer season (54.54%). In term of relative biomass consumed, in average, Himalayan tahr was the most important prey species contributed 26.27% of the biomass consumed. This was followed by yak (22.13%), cow (21.06%), musk deer (11.32%), horse (10.53%), wooly hare (1.09%), rat (0.29%), pika (0.14%) and shrew (0.07%). In average, domestic livestock including dog were contributed more biomass in the diet of snow leopard comprising 60.8% of the biomass consumed whilst the wild life species comprising 39.19%. The annual prey consumption by a snow leopard (based on 2 kg/day) was estimated to be three Himalayan tahr, seven musk deer, five wooly hare, four rat sp., two pika, one shrew and four livestock. In the present study, the highest frequency of attack was found during April to June and lowest to July to November. The day of rainy and cloudy was the more vulnerable to livestock depredation. Snow leopard attacks occurred were the highest at near escape cover such as shrub land and cliff. Both predation pressure on tahr and that on livestock suggest that the development of effective conservation strategies for two threatened species (predator and prey) depends on resolving conflicts between people and predators. Recently, direct control of free – ranging livestock, good husbandry and compensation to shepherds may reduce snow leopard – human conflict. In long term solution, the reintroduction of blue sheep at the higher altitudes could also “buffer” predation on livestock.  
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  Notes Project funded by Snow Leopard Network's Snow Leopard Conservation Grant Program. Forum of Natural Resource Managers, Nepal. Approved no  
  Call Number SLN @ rana @ 1076 Serial 887  
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Author Shrestha, R.; Wegge, P. url 
  Title Habitat relationships between wild and domestic herbivores in Nepalese trans – Himalaya Type Journal Article
  Year 2008 Publication Journal of Arid Environments Abbreviated Journal  
  Volume 72 Issue Pages 914-925  
  Keywords blue sheep; Competition; domestic; habitat partitioning; naur; Nepal; pastoralism; pseudois nayaur; trans-himalaya  
  Abstract In the semi-arid ecosystems of Asia, where pastoralism is a main subsistence occupation, grazing competition from domestic stock is believed to displace the wild ungulates. We studied the habitat relationships among sympatric naur and domestic yak and smallstock in Phu valley in upper Manang district, Nepal, on the basis of their distribution on vegetation types, elevation and slope. To control for the disturbance effect by humans, we collected the data on naur from those ranges where domestic stock were not being attended by herders. We applied correspondence analysis to explore habitat associations among animal groups (n ¬ 1415) within and across-seasons. Within each association, interspecific habitat overlaps and species habitat preferences were calculated. Naur was strongly associated with free-ranging yak as they used similar altitudinal ranges in all seasons, except in spring. Their distributions on vegetation types and slopes were also quite similar, except for a stronger preference for alpine meadows by naur during summer and winter. Naur and smallstock did not form temporal associations as the latter consistently used lower elevations. In autumn and spring, however, naur spatially overlapped with the summer range of smallstock, and both preferred the alpine meadow habitat during these periods. Alpine meadow was the least abundant vegetation type but was consistently and preferentially used by all animal groups across seasons. At high stocking densities, all three animals groups are therefore likely to compete for this vegetation type. The role of spatio-temporal heterogeneity for interpreting the interspecific relationships among ungulates in the semi-arid rangelands of the trans-Himalaya is discussed.  
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  Notes Approved no  
  Call Number SLN @ rana @ 937 Serial 891  
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Author Shrestha, R.; Wegge, P. url 
  Title Wild sheep and livestock in Nepal Trans-Himalaya: coexistence or competition? Type Journal Article
  Year 2008 Publication Environmental Conservation Abbreviated Journal  
  Volume 32 Issue 2 Pages 125-136  
  Keywords bharal; blue sheep; diet; food habits; mountain ungulates; Nepal; pastoralism; pseudois nayaur; rangeland management; resource partitioning  
  Abstract Excessive grazing by livestock is claimed to displace wild ungulates in the Trans-Himalaya. This study compares the seasonal diets and habitat use of sympatric wild naur Pseudois nayaur and domestic goat Capra hircus, sheep Ovis aries and free-ranging yak Bos grunniens in north Nepal and analyses their overlap both within and across seasons. Alpinemeadow and the legumes Oxytropis and Chesneya were critical resources for all animal groups. High overlap occurred cross-seasonally when smallstock (sheep and goats) in summer used the spring and autumn ranges of naur. Relatively high total ungulate biomass (3028 kg km-2) and low recruitment of naur (56 young per 100 adult females in autumn) suggested interspecific competition. The spatio-temporal heterogeneity in composition and phenology of food plants across the steep gradient of altitude, together with rotational grazing, appears to indirectly facilitate coexistence of naur and smallstock. However, owing to high crossseasonal (inter-seasonal) overlaps, competition is likely to occur between these two groups at high stocking densities. Within seasons, naur overlapped more with free-ranging yak than with smallstock. As their habitat use and diets were most similar in winter, when both fed extensively on the same species of shrubs, naur was most likely to compete with yak during that season.




 
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  Series Editor Series Title Abbreviated Series Title  
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  Area Expedition Conference  
  Notes Approved no  
  Call Number SLN @ rana @ 936 Serial 890  
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Author Shrestha, R.; Wegge, P.; Koirala, R.A. url 
  Title Summer diets of wild and domestic ungulates in Nepal Himalaya Type Journal Article
  Year 2005 Publication Journal of Zoology Abbreviated Journal  
  Volume 266 Issue Pages 111-119  
  Keywords argali; blue sheep; Capra hircus; diet; faecal analysis; fecal analysis; mountain ungulates; Ovis ammon hodgsoni; pseudois nayaur  
  Abstract The selection of summer forage by three sympatric ungulates in the Damodar Kunda region of upper Mustang in

north Nepal was studied to assess the extent of food overlap between them. To compare their diets, a microhistological technique of faecal analysis was used, adjusted for inherent biases by comparing it with bite-count data obtained in domestic goats. Tibetan argali Ovis ammon hodgsoni, naur (blue sheep or bharal) Pseudois nayaur and domestic goat Capra hircus consumed mostly forbs, graminoids and browse, respectively. The proportions of food items in their diets were significantly different both at the plant species (P<0.02) and at the forage category level (P<0.001). Except for sharing three common plants (Agrostis sp., Stipa sp. and Potentilla fruticosa), dietary overlap at the species level was quite low. At the forage category level, naur and domestic goat overlapped more than the other ungulate pairs. Although all three species were opportunistic, mixed feeders, argali was a more selective forb specialist grazer than the other two ungulates. Owing to some spatial separation and little dietary overlap, interspecific competition for summer forage was low. If animal densities increase, however, goats are expected to compete more with naur than with argali because of their more similar diets. Owing to differences in forage selection by argali and naur throughout their large geographical ranges, reflecting adaptations to local ecological conditions, inferences regarding forage competition between domestic livestock and these two wild caprins need to be made from local, site-specific studies, rather than from general diet comparisons.
 
  Address  
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  Publisher The Zoological Society of London Place of Publication London Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
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  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number SLN @ rana @ 934 Serial 888  
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Author Suryawanshi, K., K. url 
  Title Sunshine and the Shadow Type Magazine Article
  Year 2011 Publication Hornbill Abbreviated Journal  
  Volume Issue April-June Pages 34-37  
  Keywords Spiti, Himachal Pradesh, India, snow leopard, sighting, observation, blue sheep, Pseudois nayaur  
  Abstract Kulbhushansingh Suryawanshi shares an update on his blog which describes snow leopard sightings in Spiti, Himachal Pradesh, while studying the foraging behavior and eating habits of blue sheep (Pseudois nayaur).  
  Address  
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  Publisher Place of Publication Editor  
  Language English Summary Language Original Title  
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  Area Expedition Conference  
  Notes Approved no  
  Call Number SLN @ rana @ Serial 1360  
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Author Thapa, K. url 
  Title Is their any correlation between abundance of blue sheep population and livestock depredation by snow leopards in the Phu Valley, Manang District, Annapurna Conservation Area? Final report Type Report
  Year 2005 Publication Abbreviated Journal  
  Volume Issue Pages 1-19  
  Keywords abundance; blue; blue sheep; blue-sheep; sheep; population; livestock; livestock depredation; livestock-depredation; depredation; snow; snow leopards; snow leopard; snow-leopards; snow-leopard; leopards; leopard; valley; Manang; annapurna; annapurna conservation area; Annapurna-Conservation-Area; conservation; area; Report; project; International; international snow leopard trust; International-Snow-Leopard-Trust; trust; program; Nepal  
  Abstract This study was undertaken in the Phu valley of Manang district in the Annapurna Conservation Area, Nepal,

Spring, 2004 and 2005. I used the Snow Leopard Management Information System (“second order” survey technique), to determine

the relative abundance of snow leopards in delineated areas in Phu valley. Transects routes were plotted by

randomly selected feasible landforms such as along ridgelines, cliff bases and river bluffs where snow

leopards sign is likely to be found. Altogether, 16 transects (total length of 7.912 km) were laid down (mean

transect length=0.495 km). They revealed, 54 sign sites (both relic and non-relic) and altogether 88 signs (72

scrapes, 11 feces, 3 scent mark, 2 pugmarks and 1 hair) were recorded (6.8 site/km and 11.1 signs/km). There

were 61.1% non-relic and 38.9% relic sites. The density of snow leopards in Phu Valley may be 4-5 snow

leopards/100 kmý.It was found that the Ghyo block had the highest sign density (13.6 mean sign item/km)

and Phu block (9.8 mean sign item/km) and the lowest in Ngoru block (3.9 mean sign item/km.). For blue sheep, direct count method was applied from different appropriate vantage points (fixed-point

count). I counted total individuals in each herd and classified all individuals whenever possible, using 8 X24

binocular and 15-60x spotting scope. A total 37 blue sheep herds and 1209 individuals were observed in

192.25 kmý of the study area (blue sheep density, 6.3 kmý). Average herd size was 32.68. Herd size varied

from 1 to 103 animals (the largest so far recorded). The average sex ratio male to female for the entire survey

area was 0.67. Recruitment rate was 47.13. The ratio of yearlings to adult female was 0.45. In Ghyo block

had total 168 blue sheep (area, 44.08 km2 or 3.8/ km2 i.e. 137.2 kg/ kmý). Blue sheep density in Ngoru block

showed 4.7/km2 (area, 65.47 km2). Highest density of blue sheep among three blocks was recorded in Phu

block, 8.9/km2 (or 320 kg/km2) in its 82.70 km2 area. A standard questionnaire was designed, and interviews conducted for relevant information was collected on

livestock depredation patterns (total household survey). Out of 33 households surveyed, 30 reported that they

had livestock depredation by the snow leopard in 2004. Altogether 58 animals were reportedly lost to snow

leopards (3.1% of the total mortality). Out of the estimated standing available biomass (1, 83,483kg) in the

Phu valley at least 2220 kg or 1.3% of the total livestock biomass was consumed by snow leopards in the

year of our study (2004). It was estimated that in the Phu valley annually 1.8 animals were lost per household

to snow leopards. This means approx. Rs.413560 (US$ 5,908) is lost annually in the valley (US$

179/household/annum). Ghyo block, had the highest animals loss (53.4%), followed by Phu block (36.2%)

and Ngoru block (10.3%) to snow leopards. There is positive correlation among the densities of blue sheep, relative abundance of the snow leopard and

livestock depredation. Blue sheep is the main prey species of the snow leopard in Phu valley and its

conservation therefore matters to reduce livestock depredation. A general patterns appears here that shows

that blue sheep (prey) abundance determine snow leopard (predator) abundance and that livestock

depredation by snow leopards may be minimal where there is good population of blue sheep, and vice versa.
 
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  Area Expedition Conference  
  Notes Project funded by International Snow Leopard Trust Small Grants Program, 2005. Annapurna Conservation Area Project, Pokhara, Nepal. Approved no  
  Call Number SLN @ rana @ 1078 Serial 959  
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